:root {
  --paper: #fafaf6;
  --white: #fff;
  --forest: #153f37;
  --lagoon: #087f8c;
  --lagoon-text: #076f7a;
  --mist: #e7f4f2;
  --lagoon-wash: #dcefeb;
  --coral-wash: #f9e9e2;
  --sand: #efe6d7;
  --coral: #dd806c;
  --ink: #203c37;
  --muted: #52635e;
  --line: #d6ded7;
  --font: Arial, Helvetica, sans-serif;
  --heading: var(--font);
  --body: var(--font);
  --nav: var(--font);
  --wrap: 1200px;
}

*, *::before, *::after { box-sizing: border-box; }
html { scroll-behavior: smooth; scroll-padding-top: 112px; }
body {
  margin: 0;
  background: var(--paper);
  color: var(--ink);
  font: 400 19px/1.55 var(--body);
  -webkit-font-smoothing: antialiased;
}
body, button, input, textarea { font-family: var(--body); }
a { color: inherit; text-decoration: none; }
button, input, textarea { font: inherit; }
button { color: inherit; cursor: pointer; }
button, a { -webkit-tap-highlight-color: transparent; }
a:focus-visible, button:focus-visible, summary:focus-visible,
[tabindex]:focus-visible {
  outline: 2px solid var(--lagoon-text);
  outline-offset: 5px;
}
::selection { background: #c3e3dd; color: var(--forest); }
img, svg { display: block; max-width: 100%; }
h1, h2, h3, p, figure, dl, dd { margin: 0; }
h1, h2, h3 { color: var(--forest); overflow-wrap: break-word; }
h1, h2 {
  font-family: var(--heading);
  font-weight: 400;
  letter-spacing: -.018em;
  line-height: 1.12;
  text-wrap: balance;
}
h1 { font-size: clamp(38px, 4.6vw, 60px); }
h2 { font-size: clamp(31px, 3.2vw, 42px); }
h3 {
  font-family: var(--heading);
  font-size: 26px;
  font-weight: 400;
  line-height: 1.22;
  letter-spacing: -.012em;
}
p + p { margin-top: 1em; }
[hidden] { display: none !important; }
.wrap {
  width: min(calc(100% - 96px), var(--wrap));
  margin-inline: auto;
  min-width: 0;
}
.skip {
  position: fixed;
  z-index: 100;
  top: -100px;
  left: 20px;
  background: var(--forest);
  color: var(--white);
  padding: 10px 18px;
}
.skip:focus { top: 10px; }
.eyebrow {
  color: var(--lagoon-text);
  font: 600 11px/1.5 var(--nav);
  letter-spacing: .06em;
  text-transform: uppercase;
}
.eyebrow::before { content: none; }
.button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 18px;
  min-height: 48px;
  padding: 13px 20px;
  border: 1px solid var(--forest);
  border-radius: 6px;
  background: var(--forest);
  color: var(--white);
  font: 500 13px/1.4 var(--nav);
  text-align: center;
  transition: background-color .16s, border-color .16s;
}
.button:hover { background: #28564b; border-color: #28564b; }
.button svg { flex: 0 0 16px; width: 16px; height: 16px; transition: transform .18s ease; }
.button:hover svg, .text-link:hover svg { transform: translateX(3px); }
.button.light { background: var(--paper); border-color: var(--paper); color: var(--forest); }
.button.light:hover { background: var(--sand); border-color: var(--sand); }
.text-link {
  display: inline-flex;
  align-items: center;
  gap: 10px;
  color: var(--lagoon-text);
  padding-block: 9px;
  font: 500 13px/1.5 var(--nav);
  text-decoration: underline;
  text-decoration-thickness: 1px;
  text-underline-offset: 5px;
}
.text-link svg { flex: 0 0 16px; width: 16px; height: 16px; transition: transform .18s ease; }
.text-link:hover { color: var(--forest); }
button.text-link { border: 0; border-radius: 0; background: none; padding-inline: 0; }

/* The original wordmark remains the identity, without a replacement symbol. */
.site-header {
  position: sticky;
  top: 0;
  z-index: 30;
  border-bottom: 1px solid #b8cec6;
  border-top: 3px solid var(--lagoon);
  background: var(--paper);
}
.header-inner {
  position: relative;
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 30px;
  min-height: 94px;
  padding-block: 6px;
}
.brand { display: block; flex: 0 0 auto; }
.brand img { display: block; width: 190px; height: auto; }
.nav { display: flex; align-items: center; flex-wrap: wrap; gap: 29px; }
.nav > a:not(.button) {
  font: 500 13px/1.5 var(--nav);
  padding-block: 12px;
  text-underline-offset: 6px;
}
.nav > a:hover, .nav > a[aria-current] { color: var(--lagoon-text); text-decoration: underline; text-decoration-color: var(--coral); }
.nav .button { min-height: 44px; padding: 10px 15px; font-size: 12px; gap: 12px; }
.nav-services { position: relative; font: 500 13px/1.5 var(--nav); }
.nav-services > summary { padding-block: 12px; list-style: none; cursor: pointer; text-underline-offset: 6px; }
.nav-services > summary::-webkit-details-marker { display: none; }
.nav-services > summary::after { content: '+'; margin-left: 8px; color: var(--lagoon-text); }
.nav-services[open] > summary::after { content: '−'; }
.nav-services[data-active] > summary, .nav-services > summary:hover { color: var(--lagoon-text); text-decoration: underline; text-decoration-color: var(--coral); }
.service-menu { position: absolute; right: 0; top: 100%; z-index: 35; width: 282px; padding: 8px; background: var(--paper); border: 1px solid var(--line); box-shadow: 0 12px 26px #153f3712; }
.service-menu a { display: block; min-height: 44px; padding: 12px; color: var(--forest); font-size: 14px; }
.service-menu a:hover, .service-menu a[aria-current] { background: var(--lagoon-wash); text-decoration: underline; text-underline-offset: 4px; }
.menu-toggle {
  display: none;
  width: 44px;
  height: 44px;
  align-items: center;
  justify-content: center;
  padding: 10px;
  background: none;
  border: 1px solid #a9bbb1;
  border-radius: 5px;
}
.menu-toggle svg { width: 22px; height: 22px; }

/* The introduction is led by the pathologist and the scope of the work. */
.hero { padding-block: 64px 68px; }
.hero-personal > .eyebrow { margin-bottom: 24px; }
.hero-personal h1 { max-width: 1120px; font-size: clamp(46px, 5.6vw, 76px); line-height: 1.07; letter-spacing: -.025em; }
.hero-personal h1 span { display: block; color: var(--lagoon-text); }
.hero-lower { display: grid; grid-template-columns: minmax(0, 1.55fr) minmax(0, .8fr); gap: 100px; align-items: start; margin-top: 42px; }
.hero-lower > * { min-width: 0; }
.hero-copy {
  max-width: 620px;
  color: var(--muted);
  font-size: 23px;
  line-height: 1.5;
}
.hero-actions { display: flex; align-items: center; flex-wrap: wrap; gap: 25px; margin-top: 28px; }
.hero-context { padding-left: 28px; border-left: 1px solid var(--lagoon); }
.hero-context > .eyebrow { margin-bottom: 17px; }
.hero-context > p:not(.eyebrow) { color: var(--muted); font-size: 18px; line-height: 1.5; }
.hero-facts { display: grid; gap: 14px; }
.hero-facts > div { display: grid; grid-template-columns: 88px minmax(0, 1fr); gap: 12px; align-items: baseline; }
.hero-facts dt { color: var(--muted); font: 500 10px/1.5 var(--nav); }
.hero-facts dd { color: var(--forest); font-size: 17px; line-height: 1.4; }
.hero-context > p:not(.eyebrow) + .hero-facts { margin-top: 20px; }

.section { padding-block: 72px; }
#services { background: var(--mist); border-block: 1px solid #c1ddd5; }
.section-heading { display: flex; justify-content: space-between; align-items: end; gap: 70px; margin-bottom: 37px; }
.section-heading h2 { max-width: 670px; margin-top: 15px; }
.section-heading > p { max-width: 370px; color: var(--muted); font-size: 20px; }
.services-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 43px; }
.service-card { display: flex; flex-direction: column; align-items: start; padding-top: 24px; border-top: 4px solid var(--lagoon); min-width: 0; }
.service-card[data-service='toxicologic'] { border-color: var(--lagoon); }
.service-card[data-service='investigative'] { border-color: var(--forest); }
.service-card[data-service='computational'] { border-color: var(--coral); }
.service-card h3 { margin-bottom: 16px; font-size: 27px; }
.service-card h3 a:hover { text-decoration: underline; text-underline-offset: 5px; }
.service-card p { color: var(--muted); font-size: 19px; }
.service-card .service-output { margin-top: 16px; font-size: 17px; color: var(--forest); }
.service-card .text-link { margin-top: auto; padding-top: 22px; font-size: 12px; }
.service-top, .service-index, .service-icon { display: none; }
.service-row { display: grid; grid-template-columns: 48px minmax(0, .95fr) minmax(0, 1.15fr); gap: 38px; padding-block: 31px 34px; border-top: 1px solid var(--lagoon); }
.service-row[data-service='investigative'] { border-top-color: var(--forest); }
.service-row[data-service='computational'] { border-top-color: var(--coral); }
.service-row:last-child { padding-bottom: 0; }
.service-row > * { min-width: 0; }
.service-number { padding-top: 7px; color: var(--lagoon-text); font: 500 11px/1.5 var(--nav); }
.service-row h3 { max-width: 420px; font-size: clamp(26px, 2.5vw, 34px); line-height: 1.16; }
.service-row h3 a:hover { color: var(--lagoon-text); }
.service-summary > p { max-width: 550px; color: var(--muted); font-size: 20px; line-height: 1.5; }
.service-summary .text-link { margin-top: 13px; }

.experience-preview { background: #f5eee2; border-bottom: 1px solid #e4d8c5; }
.featured-work { display: grid; grid-template-columns: minmax(0, 1.05fr) minmax(0, .95fr); gap: 52px; align-items: center; padding: 36px; border: 1px solid #d3c6b3; border-radius: 8px; background: #fbf9f2; }
.featured-work > * { min-width: 0; }
.featured-work h3 { max-width: 600px; margin-block: 15px 20px; font-size: clamp(28px, 2.8vw, 36px); line-height: 1.16; }
.featured-work-copy > p:not(.eyebrow) { color: var(--muted); font-size: 20px; line-height: 1.55; }
.featured-work .text-link { margin-top: 17px; }
.featured-work .histology-placeholder { width: 100%; max-width: none; margin: 0; }
.featured-work .placeholder-art { min-height: 220px; max-height: 320px; }
.work-notes { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 54px; margin-top: 35px; }
.work-note { min-width: 0; padding-top: 24px; border-top: 1px solid #93b4a9; }
.work-note:nth-child(2) { border-top-color: var(--coral); }
.work-note h3 { margin-block: 14px 16px; font-size: 26px; }
.work-note > p:not(.eyebrow) { color: var(--muted); font-size: 19px; }
.work-note .text-link { margin-top: 15px; }
.work-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 43px; }
.work-item { border-top: 1px solid #a7bfb3; padding-top: 23px; min-width: 0; }
.work-item:nth-child(1) { border-top-color: var(--lagoon); }
.work-item:nth-child(2) { border-top-color: var(--forest); }
.work-item:nth-child(3) { border-top-color: var(--coral); }
.work-category { margin-bottom: 14px; color: var(--lagoon-text); font: 600 10px/1.5 var(--nav); letter-spacing: .08em; text-transform: uppercase; }
.work-item h3 { margin-bottom: 15px; font-size: 26px; }
.work-item p { font-size: 19px; color: var(--muted); }
.work-tags, .method-tags { color: var(--muted); font-size: 15px; line-height: 1.65; margin-top: 17px; }
.work-tags span, .method-tags span { display: inline; }
.work-tags span + span::before, .method-tags span + span::before { content: ' / '; white-space: pre; color: #627c70; }

.about-grid { display: grid; grid-template-columns: minmax(0, .8fr) minmax(0, 1.2fr); gap: 105px; align-items: start; }
.about-copy h2 { margin-block: 15px 24px; max-width: 660px; }
.about-copy p { color: var(--muted); font-size: 20px; }
.about-copy .text-link { margin-top: 17px; }
.personal-intro { display: grid; grid-template-columns: minmax(0, .75fr) minmax(0, 1.25fr); gap: 100px; align-items: start; }
.personal-meta, .personal-copy { min-width: 0; }
.personal-meta { padding-top: 20px; border-top: 1px solid var(--coral); }
.personal-meta h2 { margin-top: 16px; font-size: clamp(30px, 3vw, 40px); }
.personal-copy > p { color: var(--muted); font-size: 21px; line-height: 1.6; }
.personal-copy .related-services { margin-top: 20px; }
.about-identity { padding-left: 31px; border-left: 3px solid var(--coral); min-width: 0; }
.about-identity h3 { font-size: 28px; margin-bottom: 10px; }
.about-identity .name { font: 400 28px/1.2 var(--heading); color: var(--forest); }
.credential { margin-top: 8px; color: var(--muted); font-size: 17px; line-height: 1.5; }
.career-list { margin-top: 26px; }
.career-list > div { padding-block: 15px; border-top: 1px solid var(--line); }
.career-list dt { color: var(--forest); font: 600 13px/1.5 var(--nav); }
.career-list dd { margin-top: 4px; color: var(--muted); font-size: 17px; line-height: 1.4; }
.working-note { max-width: 900px; padding-block: 20px 72px; color: var(--muted); font-size: 20px; line-height: 1.6; }
.working-note h2 { margin-bottom: 20px; }
.working-note strong { color: var(--forest); font-weight: 600; }
.working-note > .text-link { margin-top: 22px; }

/* Working arrangements pair each assignment with its practical scope. */
.engagement-row { display: grid; grid-template-columns: minmax(0, .95fr) minmax(0, 1.05fr); gap: 85px; padding-block: 40px 46px; border-top: 1px solid var(--lagoon); }
.engagement-row:nth-of-type(3n + 2) { border-top-color: var(--forest); }
.engagement-row:nth-of-type(3n) { border-top-color: var(--coral); }
.engagement-summary, .engagement-details { min-width: 0; }
.engagement-summary .eyebrow { margin-bottom: 14px; }
.engagement-summary h2 { margin-block: 0 22px; }
.engagement-summary > p:not(.eyebrow) { max-width: 550px; color: var(--muted); font-size: 20px; line-height: 1.55; }
.engagement-summary > .text-link { margin-top: 17px; }
.engagement-details .measurement-list { margin-top: 0; }
.engagement-details .measurement-list > div:first-child { padding-top: 0; border-top: 0; }
.engagement-details > .text-link, .engagement-details > .button { margin-top: 17px; }
.engagement-process { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 34px 65px; margin: 32px 0 0; padding-left: 24px; }
.engagement-process li { min-width: 0; padding-left: 8px; }
.engagement-process li::marker { color: var(--lagoon-text); font: 600 13px/1.5 var(--nav); }
.engagement-process h3 { margin-bottom: 13px; font-size: 26px; }
.engagement-process p { color: var(--muted); font-size: 20px; line-height: 1.55; }

.contact-band { background: var(--forest); color: var(--paper); border-top: 3px solid var(--coral); }
.contact-inner { display: grid; grid-template-columns: minmax(0, 1fr) minmax(0, .68fr); align-items: center; gap: 80px; padding-block: 57px; }
.contact-band h2 { margin-top: 15px; font-size: 42px; color: var(--paper); }
.contact-band p { max-width: 610px; margin-top: 20px; font-size: 21px; color: #d5e6dc; }
.contact-band .eyebrow { color: #a6dcda; margin-top: 0; font-size: 11px; }
.contact-band .email-link { color: var(--paper); text-decoration-color: #93bcae; }
.contact-band .email-link:hover { color: #bce2df; }
.contact-band a:focus-visible { outline-color: var(--sand); }
.contact-actions { display: flex; flex-direction: column; align-items: start; justify-self: end; gap: 20px; min-width: 0; }
.email-link { color: var(--forest); font: 400 20px/1.45 var(--body); text-decoration: underline; text-decoration-thickness: 1px; text-underline-offset: 5px; overflow-wrap: anywhere; }
.email-link:hover { color: var(--lagoon-text); }
.site-footer { padding-block: 32px 23px; }
.footer-top { display: flex; justify-content: space-between; align-items: center; flex-wrap: wrap; gap: 24px; }
.site-footer .brand img { width: 175px; }
.footer-top p { max-width: 340px; font-size: 17px; color: var(--muted); }
.footer-links { display: flex; flex-wrap: wrap; gap: 24px; font: 500 12px/1.5 var(--nav); }
.footer-links a:hover { text-decoration: underline; text-underline-offset: 4px; }
.footer-bottom { display: flex; justify-content: space-between; gap: 24px; margin-top: 27px; padding-top: 19px; border-top: 1px solid var(--line); color: var(--muted); font-size: 13px; }
.footer-bottom span:last-child { text-align: right; }

/* Detail pages use the same typographic rhythm, without decorative panels. */
.subhero { padding-block: 28px 53px; }
.breadcrumb { display: flex; align-items: center; flex-wrap: wrap; gap: 10px; margin-bottom: 41px; color: var(--muted); font-size: 14px; line-height: 1.5; }
.breadcrumb a:hover { text-decoration: underline; }
.subhero h1 { max-width: 980px; margin-top: 19px; font-size: clamp(38px, 4.6vw, 60px); line-height: 1.1; letter-spacing: -.02em; }
.subhero .lead { max-width: 840px; margin-top: 24px; font-size: 24px; line-height: 1.5; color: var(--muted); }
.service-page-top { max-width: 970px; }
.service-page-top h1 { max-width: 960px; }
.service-page-top .lead { max-width: 830px; }
.subsection { padding-block: 59px; border-top: 1px solid var(--line); }
.subsection h2 { margin-top: 15px; }
.service-intro { max-width: 940px; color: var(--ink); font-size: 23px; line-height: 1.6; }
.project-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 43px; margin-top: 35px; }
.project-item { min-width: 0; padding-top: 22px; border-top: 1px solid #9cb4a7; }
.project-item h3 { font-size: 25px; margin-bottom: 15px; }
.project-item p { font-size: 19px; color: var(--muted); }
.output-section { background: var(--lagoon-wash); border-block: 1px solid #c1ddd5; }
.output-grid { display: grid; grid-template-columns: minmax(0, .9fr) minmax(0, 1.1fr); gap: 90px; }
.output-grid h2 { margin-top: 16px; }
.output-grid .intro { color: var(--muted); font-size: 20px; margin-top: 20px; }
.output-list { list-style: none; padding: 0; margin: 0; }
.output-list li { border-top: 1px solid #afc6ba; padding-block: 18px; font-size: 21px; line-height: 1.5; }
.scope-note { color: var(--muted); margin-top: 22px; font-size: 16px; }
.scope-grid { display: grid; grid-template-columns: minmax(0, .9fr) minmax(0, 1.1fr); gap: 90px; }
.scope-questions { margin: 0; padding-left: 24px; }
.scope-questions li { padding-left: 7px; margin-bottom: 19px; font-size: 21px; }
.scope-questions li::marker { color: var(--lagoon-text); font-size: 16px; }
.specialism { max-width: 950px; }
.specialism h2, .specialism h3 { margin-bottom: 23px; }
.specialism p, .specialism li { color: var(--muted); font-size: 20px; }
.specialism ul { padding-left: 24px; margin-top: 23px; }
.specialism li { padding-left: 5px; margin-block: 9px; }
.related-services { display: flex; align-items: start; flex-wrap: wrap; gap: 20px 35px; margin-top: 24px; }
.about-page-grid { display: grid; grid-template-columns: minmax(0, .72fr) minmax(0, 1.28fr); gap: 90px; align-items: start; }
.about-page-grid .about-identity { border-left: 0; border-top: 1px solid #cbbda9; padding: 24px 0 0; }
.about-page-text h2 { margin-top: 0; }
.about-page-text p { font-size: 21px; color: var(--muted); }

.contact-page { max-width: 1040px; padding-block: 28px 73px; }
.contact-page h1 { margin-top: 19px; font-size: clamp(42px, 4.3vw, 60px); }
.contact-page .scope-note + h2 { margin-top: 32px; }
.contact-page .lead { max-width: 730px; margin-top: 26px; color: var(--muted); font-size: 24px; line-height: 1.5; }
.contact-email-block { display: flex; justify-content: space-between; align-items: center; flex-wrap: wrap; gap: 25px 35px; border-block: 1px solid #afc4b7; padding-block: 31px; margin-block: 39px; }
.contact-email-block .email-link { font-size: 29px; }
.contact-email-block .eyebrow { margin-bottom: 11px; }
.contact-email-block > div { min-width: 0; }
.contact-checklist { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 42px; margin-top: 33px; }
.contact-checklist h3 { font-size: 25px; margin-bottom: 13px; }
.contact-checklist p { color: var(--muted); font-size: 19px; }

.experience-section { padding-block: 58px; border-top: 1px solid var(--line); }
.experience-intro { max-width: 890px; color: var(--muted); font-size: 23px; line-height: 1.6; }
.experience-section .experience-analysis-heading > p { max-width: none; }
.experience-section .experience-analysis-heading { flex-wrap: wrap; column-gap: 40px; row-gap: 16px; }
@media (min-width: 760px) {
  .experience-analysis-heading > p { white-space: nowrap; }
}
.experience-meta > .editorial-illustration { margin-top: 20px; }
.experience-list { margin-top: 39px; }
.experience-row { display: grid; grid-template-columns: minmax(0, .65fr) minmax(0, 1.35fr); gap: 70px; padding-block: 30px 33px; border-top: 1px solid #afc4b7; }
.experience-meta { color: var(--muted); font-size: 17px; line-height: 1.5; }
.experience-meta small { display: block; color: var(--lagoon-text); font: 600 10px/1.6 var(--nav); letter-spacing: .09em; text-transform: uppercase; }
.experience-meta p, .experience-meta strong { display: block; margin-top: 9px; color: var(--forest); font-weight: 400; }
.experience-detail h3 { max-width: 780px; margin-bottom: 17px; font-size: 28px; }
.experience-detail p { color: var(--muted); font-size: 20px; }
.experience-detail .method-tags { font-size: 15px; line-height: 1.65; }
.work-item .work-category { font-size: 10px; color: var(--lagoon-text); }
.publication-note { max-width: 940px; border-top: 1px solid var(--line); padding-top: 23px; color: var(--muted); font-size: 18px; }
.publication-note a { color: var(--lagoon-text); text-decoration: underline; text-underline-offset: 4px; }

/* Toxicologic pathology follows the distinct roles and study requirements. */
[id] { scroll-margin-top: 8px; }
.tox-overview { max-width: 950px; color: var(--ink); font-size: 23px; line-height: 1.6; }
.toc-links { display: flex; flex-wrap: wrap; align-items: center; gap: 10px 29px; padding-block: 21px; margin-top: 25px; border-block: 1px solid var(--line); }
.toc-links a { color: var(--lagoon-text); font: 500 13px/1.5 var(--nav); text-decoration: underline; text-underline-offset: 5px; }
.toc-links a:hover { color: var(--forest); }
.tox-role-grid { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 58px; padding-block: 55px; }
.tox-role { display: flex; flex-direction: column; align-items: start; min-width: 0; padding-top: 26px; border-top: 2px solid var(--lagoon); }
.tox-role:nth-child(2) { border-top-color: var(--forest); }
.tox-role .eyebrow { margin-bottom: 14px; }
.tox-role h2 { margin-bottom: 23px; font-size: clamp(31px, 3vw, 40px); }
.tox-role p, .tox-role li { color: var(--muted); font-size: 20px; line-height: 1.55; }
.tox-role ul { margin: 22px 0 25px; padding-left: 22px; }
.tox-role li { padding-left: 5px; }
.tox-role li + li { margin-top: 11px; }
.tox-role li::marker { color: var(--lagoon-text); }
.tox-role .histology-placeholder { width: 100%; max-width: none; margin-top: auto; }
.study-grid { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 37px 55px; margin-top: 35px; }
.study-grid > article { min-width: 0; padding-top: 22px; border-top: 2px solid var(--lagoon); }
.study-grid > article:nth-child(2n) { border-top-color: var(--coral); }
.study-grid h3 { margin-bottom: 15px; font-size: 27px; }
.study-grid p { color: var(--muted); font-size: 20px; }
.planning-grid, .quality-grid, .delivery-grid { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 65px; align-items: start; }
.planning-grid > *, .quality-grid > *, .delivery-grid > * { min-width: 0; }
.planning-grid h2, .quality-grid h2, .delivery-grid h2, .service-heading { margin-bottom: 23px; }
.planning-grid p, .planning-grid li, .quality-grid p, .quality-grid li, .delivery-grid p, .delivery-grid li { color: var(--muted); font-size: 20px; }
.planning-grid ul, .quality-grid ul, .delivery-grid ul { padding-left: 24px; margin-top: 23px; }
.planning-grid li + li, .quality-grid li + li, .delivery-grid li + li { margin-top: 11px; }
.planning-grid .histology-placeholder, .delivery-grid .histology-placeholder { margin-top: 0; width: 100%; max-width: none; }
.quality-grid .measurement-list { margin-top: 0; }
.quality-note { max-width: 950px; margin-top: 26px; color: var(--muted); font-size: 17px; line-height: 1.6; }
.image-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 27px; margin-top: 35px; }
.image-grid > * { min-width: 0; }
.image-grid .histology-placeholder { width: 100%; max-width: none; margin: 0; }
.image-grid .placeholder-art { min-height: 150px; max-height: 245px; }
.image-grid .histology-placeholder:nth-child(2) .placeholder-art { background: #e1e9dd; }
.image-grid .histology-placeholder:nth-child(3) .placeholder-art { background: var(--coral-wash); color: #805447; }
.personal-portrait { width: 100%; max-width: 340px; }
.personal-portrait img { width: 100%; height: auto; aspect-ratio: 3 / 4; border-radius: 6px; }
.personal-portrait figcaption { margin-top: 9px; color: var(--muted); font-size: 13px; line-height: 1.45; }
.personal-meta .personal-portrait { max-width: 280px; margin-top: 27px; }

/* Image spaces are deliberately unfilled and labelled for later histology. */
.histology-placeholder { margin: 27px 0 0; max-width: 540px; min-width: 0; border: 1px dashed #89ada1; border-radius: 6px; overflow: hidden; background: #f0f7f2; }
.placeholder-art { width: 100%; display: flex; flex-direction: column; align-items: center; justify-content: center; gap: 14px; min-height: 170px; aspect-ratio: 4 / 3; max-height: 290px; padding: 22px; color: #42685c; background: #e4f1eb; }
.placeholder-art svg { width: 44px; height: 44px; opacity: .65; flex: 0 0 auto; }
.placeholder-art > span { display: block; font: 500 11px/1.5 var(--nav); letter-spacing: .08em; text-transform: uppercase; }
.histology-placeholder figcaption { padding: 14px 17px 15px; background: rgba(255,255,255,.65); font-size: 15px; line-height: 1.45; color: var(--forest); }
.histology-placeholder figcaption strong { display: block; font-weight: 600; }
.histology-placeholder figcaption > span { display: block; color: var(--muted); margin-top: 5px; font-size: 13px; }
.work-item .histology-placeholder { max-width: none; margin: 0 0 23px; }
.work-item .placeholder-art { min-height: 160px; max-height: 235px; }
.work-item:nth-child(2) .placeholder-art { background: #e1e9dd; }
.work-item:nth-child(3) .placeholder-art { background: var(--coral-wash); color: #805447; }
.experience-detail .histology-placeholder { max-width: 500px; margin-top: 28px; }
.experience-detail .histology-placeholder figcaption { font-size: 15px; }
.experience-detail .histology-placeholder figcaption > span { font-size: 13px; }

/* Histomorphometry has its own concrete, illustrated area. */
#histomorph-heading { margin-bottom: 23px; }
.histomorph-intro { max-width: 920px; color: var(--ink); font-size: 23px; line-height: 1.6; }
.histomorph-feature { display: grid; grid-template-columns: minmax(0, 1.1fr) minmax(0, .9fr); align-items: center; gap: 72px; margin-block: 58px 72px; padding: 42px; background: var(--coral-wash); border-left: 4px solid var(--coral); }
.histomorph-feature h2 { margin-block: 14px 20px; }
.histomorph-feature p { color: var(--muted); font-size: 20px; }
.tox-role .eyebrow,
.planning-grid .eyebrow,
.quality-grid .eyebrow,
.delivery-grid .eyebrow,
.specialism .eyebrow,
.histomorph-feature .eyebrow {
  color: var(--lagoon-text);
  font: 600 10px/1.5 var(--nav);
  letter-spacing: .06em;
  text-transform: uppercase;
}
.histomorph-feature .text-link { margin-top: 16px; }
.histomorph-feature .histology-placeholder { margin-top: 0; }
.histomorph-feature .placeholder-art { max-height: 250px; background: #f3e4d5; color: #665744; }
.histomorph-grid { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 48px 55px; margin-top: 42px; }
.histomorph-examples-heading { margin-top: 39px; }
.histomorph-item { min-width: 0; padding-top: 26px; border-top: 2px solid var(--lagoon); display: flex; flex-direction: column; }
.histomorph-item:nth-child(2n) { border-top-color: var(--forest); }
.histomorph-item:nth-child(3n) { border-top-color: var(--coral); }
.histomorph-item h3 { margin-bottom: 17px; font-size: 28px; }
.histomorph-item p { font-size: 20px; color: var(--muted); }
.histomorph-item .histology-placeholder { margin-top: auto; max-width: none; }
.histomorph-item .measurement-list { margin-bottom: 26px; }
.histomorph-item .placeholder-art { max-height: 275px; }
.measurement-list { margin-top: 23px; }
.measurement-list > div { border-top: 1px solid var(--line); padding-block: 15px; }
.measurement-list dt { font: 600 12px/1.5 var(--nav); color: var(--forest); }
.measurement-list dd { margin: 6px 0 0; color: var(--muted); font-size: 18px; line-height: 1.5; }
.sampling-note { margin-block: 58px; padding: 33px 36px; border-left: 4px solid var(--forest); background: var(--lagoon-wash); }
.sampling-note h2, .sampling-note h3 { margin-bottom: 18px; }
.sampling-note p, .sampling-note li { font-size: 20px; color: var(--ink); }
.sampling-note ul { padding-left: 24px; margin-bottom: 0; }
.sampling-note li + li { margin-top: 10px; }

/* The name is explained once, with its own illustrated literary reference. */
.name-story { display: grid; grid-template-columns: minmax(0, 1fr) minmax(0, 1fr); gap: 52px; align-items: center; margin-block: 0 54px; padding: 36px; background: var(--sand); border-left: 3px solid var(--coral); }
.name-story-copy, .name-story-figure { min-width: 0; }
.name-story .eyebrow { margin-bottom: 15px; }
.name-story h2 { margin-bottom: 22px; font-family: var(--heading); font-weight: 400; letter-spacing: -.025em; }
.name-story-text { color: var(--ink); font-size: 20px; line-height: 1.6; }
.name-story-source { margin-top: 20px; color: var(--muted); font-size: 14px; line-height: 1.5; }
.name-story-source a { text-decoration: underline; text-underline-offset: 3px; }
.name-story-figure img { width: 100%; height: auto; }
.name-story-figure figcaption { margin-top: 12px; color: var(--muted); font-size: 13px; line-height: 1.5; }

@media (max-width: 1050px) {
  .name-story { gap: 32px; padding: 28px; }
  .wrap { width: calc(100% - 64px); }
  .nav { gap: 21px; }
  .nav > a:not(.button) { font-size: 12px; }
  .nav .button { font-size: 11px; padding-inline: 12px; }
  .brand img { width: 185px; }
  .header-inner { min-height: 94px; gap: 25px; }
  .hero { padding-block: 56px 60px; }
  .hero-lower { gap: 60px; }
  .hero-context { padding-left: 23px; }
  .hero-facts > div { grid-template-columns: 75px minmax(0, 1fr); gap: 10px; }
  .service-row { grid-template-columns: 34px minmax(0, .95fr) minmax(0, 1.15fr); gap: 29px; }
  .featured-work { gap: 36px; padding: 28px; }
  .work-notes { gap: 38px; }
  .personal-intro { gap: 65px; }
  .services-grid, .work-grid, .project-grid { gap: 29px; }
  .service-card h3 { font-size: 25px; }
  .section-heading { gap: 40px; }
  .section-heading > p { max-width: 300px; }
  .about-grid { gap: 65px; }
  .about-page-grid, .output-grid, .scope-grid { gap: 60px; }
  .contact-inner { gap: 45px; }
  .contact-actions .email-link { font-size: 18px; }
  .experience-row { gap: 45px; }
  .engagement-row { gap: 55px; }
  .engagement-process { column-gap: 45px; }
  .histomorph-feature { gap: 40px; padding: 33px; }
  .histomorph-grid { gap: 38px; }
}

@media (max-width: 800px) {
  html { scroll-padding-top: 106px; }
  .wrap { width: calc(100% - 48px); }
  .header-inner { min-height: 90px; }
  .brand img { width: 180px; }
  .nav { gap: 17px; }
  .nav > a:not(.button) { font-size: 11px; }
  .nav .button { font-size: 10px; padding: 10px 12px; gap: 9px; }
  .hero { padding-block: 49px 54px; }
  .hero-personal h1 { font-size: 55px; }
  .hero-lower { grid-template-columns: minmax(0, 1.35fr) minmax(0, .9fr); gap: 35px; margin-top: 35px; }
  .hero-copy { font-size: 21px; }
  .hero-context { padding-left: 19px; }
  .hero-facts > div { grid-template-columns: minmax(0, 1fr); gap: 3px; }
  .hero-facts { gap: 12px; }
  .hero-actions { gap: 17px; }
  .hero-actions .button { font-size: 12px; padding-inline: 17px; }
  .hero-actions .text-link { font-size: 12px; }
  .section { padding-block: 58px; }
  .section-heading { align-items: start; gap: 30px; }
  .section-heading > p { font-size: 19px; max-width: 260px; }
  .services-grid, .work-grid, .project-grid { gap: 23px; }
  .service-card h3, .work-item h3 { font-size: 23px; }
  .service-card p, .work-item p { font-size: 18px; }
  .service-card .text-link { font-size: 11px; }
  .service-row { gap: 24px; }
  .service-row h3 { font-size: 25px; }
  .service-summary > p { font-size: 19px; }
  .featured-work { gap: 28px; padding: 24px; }
  .featured-work h3 { font-size: 29px; }
  .featured-work .placeholder-art { min-height: 210px; }
  .work-notes { gap: 30px; }
  .work-note h3 { font-size: 24px; }
  .personal-intro { gap: 42px; }
  .personal-meta h2 { font-size: 31px; }
  .personal-copy > p { font-size: 20px; }
  .about-grid, .about-page-grid, .output-grid, .scope-grid { gap: 40px; }
  .about-identity { padding-left: 23px; }
  .about-copy p { font-size: 19px; }
  .about-page-text p { font-size: 20px; }
  .contact-inner { grid-template-columns: minmax(0, 1fr); gap: 28px; padding-block: 45px; }
  .contact-actions { flex-direction: row; align-items: center; flex-wrap: wrap; gap: 22px; justify-self: start; }
  .footer-top p { max-width: 300px; }
  .footer-links { width: 100%; }
  .subhero { padding-bottom: 44px; }
  .subhero .lead { font-size: 23px; }
  .subsection, .experience-section { padding-block: 48px; }
  .project-item h3 { font-size: 23px; }
  .project-item p { font-size: 18px; }
  .experience-row { grid-template-columns: minmax(0, .58fr) minmax(0, 1.42fr); gap: 32px; }
  .experience-detail h3 { font-size: 26px; }
  .tox-role-grid { gap: 35px; }
  .planning-grid, .quality-grid, .delivery-grid { gap: 38px; }
  .study-grid { gap: 31px 38px; }
  .image-grid { grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 23px; }
  .histomorph-feature { gap: 30px; padding: 28px; margin-block: 43px 58px; }
  .histomorph-grid { gap: 32px; }
  .histomorph-item h3 { font-size: 26px; }
  .histomorph-item p { font-size: 19px; }
  .placeholder-art { min-height: 150px; }
}

@media (max-width: 680px) {
  .name-story { grid-template-columns: minmax(0, 1fr); gap: 27px; padding: 24px 22px; margin-bottom: 38px; }
  html { scroll-padding-top: 103px; }
  body { font-size: 19px; }
  .wrap { width: calc(100% - 40px); }
  .header-inner { min-height: 88px; padding-block: 6px; gap: 20px; }
  .brand img { width: 170px; }
  .menu-toggle { display: flex; flex: 0 0 44px; }
  .nav { display: none; position: absolute; top: 100%; left: -20px; right: -20px; padding: 15px 20px 23px; background: var(--paper); border-bottom: 1px solid var(--line); }
  .nav.is-open { display: flex; flex-direction: column; align-items: stretch; gap: 0; }
  .nav > a:not(.button) { padding-block: 13px; font-size: 15px; }
  .nav .button { align-self: start; min-width: 200px; margin-top: 9px; padding: 13px 17px; font-size: 12px; }
  .hero { padding-block: 39px 44px; }
  .hero-personal > .eyebrow { margin-bottom: 22px; font-size: 10px; }
  .hero-personal h1 { max-width: 590px; font-size: clamp(38px, 10.25vw, 52px); letter-spacing: -.025em; }
  .hero-lower { grid-template-columns: minmax(0, 1fr); gap: 30px; margin-top: 28px; }
  .hero-copy { max-width: 570px; font-size: 21px; }
  .hero-actions { gap: 14px 20px; margin-top: 25px; }
  .hero-actions .button { min-height: 48px; font-size: 12px; }
  .hero-actions .text-link { font-size: 12px; }
  .hero-context { padding: 21px 0 0; border-left: 0; border-top: 1px solid #9ec2b8; }
  .hero-context > .eyebrow { margin-bottom: 15px; }
  .hero-facts { grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 16px; }
  .hero-facts dd { font-size: 16px; }
  h2 { font-size: clamp(30px, 7.8vw, 36px); }
  h3 { font-size: 25px; }
  .eyebrow { font-size: 10px; }
  .section { padding-block: 49px; }
  .section-heading { display: block; margin-bottom: 29px; }
  .section-heading h2 { margin-top: 13px; }
  .section-heading > p { max-width: 100%; margin-top: 20px; font-size: 19px; }
  .services-grid, .work-grid, .project-grid { grid-template-columns: minmax(0, 1fr); gap: 28px; }
  .service-card { padding-top: 20px; }
  .service-card h3, .work-item h3 { font-size: 27px; margin-bottom: 13px; }
  .service-card p, .work-item p { font-size: 19px; }
  .service-card .service-output { font-size: 17px; margin-top: 14px; }
  .service-card .text-link { padding-top: 17px; font-size: 12px; }
  .service-row { grid-template-columns: 26px minmax(0, 1fr); gap: 18px; padding-block: 26px 29px; }
  .service-number { padding-top: 5px; }
  .service-row h3 { font-size: 27px; }
  .service-summary { grid-column: 1 / -1; }
  .service-summary > p { font-size: 19px; }
  .service-summary .text-link { margin-top: 10px; }
  .featured-work { grid-template-columns: minmax(0, 1fr); gap: 27px; padding: 24px 22px; }
  .featured-work h3 { font-size: 29px; }
  .featured-work .placeholder-art { min-height: 190px; max-height: 265px; }
  .work-notes { grid-template-columns: minmax(0, 1fr); gap: 27px; margin-top: 29px; }
  .work-note { padding-top: 22px; }
  .work-note h3 { font-size: 25px; }
  .work-item { padding-top: 20px; }
  .work-category { margin-bottom: 12px; }
  .work-tags, .method-tags { font-size: 16px; margin-top: 14px; }
  .about-grid, .about-page-grid, .output-grid, .scope-grid { grid-template-columns: minmax(0, 1fr); gap: 31px; }
  .about-copy { grid-row: 1; }
  .about-copy h2 { margin-block: 13px 21px; }
  .about-copy p { font-size: 20px; }
  .personal-intro { grid-template-columns: minmax(0, 1fr); gap: 25px; }
  .personal-meta { padding-top: 19px; }
  .personal-meta h2 { margin-top: 13px; font-size: 33px; }
  .personal-copy > p { font-size: 20px; }
  .about-identity { padding: 23px 0 0; border-left: 0; border-top: 1px solid #cbbda9; }
  .about-identity h3, .about-identity .name { font-size: 28px; }
  .credential { font-size: 18px; }
  .career-list { margin-top: 22px; }
  .career-list > div { padding-block: 13px; }
  .career-list dd { font-size: 18px; }
  .working-note { font-size: 20px; }
  .engagement-row { grid-template-columns: minmax(0, 1fr); gap: 25px; padding-block: 30px 34px; }
  .engagement-summary h2 { margin-bottom: 19px; }
  .engagement-summary > .text-link { margin-top: 13px; }
  .engagement-process { grid-template-columns: minmax(0, 1fr); gap: 26px; margin-top: 26px; }
  .engagement-process h3 { font-size: 25px; margin-bottom: 11px; }
  .contact-inner { gap: 26px; padding-block: 43px; }
  .contact-band h2 { font-size: clamp(32px, 8.5vw, 38px); }
  .contact-band p { font-size: 21px; margin-top: 19px; }
  .contact-actions { flex-direction: column; align-items: start; gap: 20px; }
  .contact-actions .email-link { font-size: 20px; }
  .site-footer { padding-block: 27px 21px; }
  .footer-top { display: block; }
  .site-footer .brand img { width: 170px; }
  .footer-top p { margin-top: 21px; max-width: 330px; font-size: 17px; }
  .footer-links { gap: 15px 24px; margin-top: 24px; font-size: 12px; }
  .footer-bottom { display: block; font-size: 13px; margin-top: 23px; }
  .footer-bottom span { display: block; }
  .footer-bottom span:last-child { margin-top: 7px; text-align: left; }
  .subhero { padding-block: 24px 39px; }
  .breadcrumb { margin-bottom: 30px; gap: 7px; font-size: 13px; }
  .subhero h1 { margin-top: 17px; font-size: clamp(34px, 8.7vw, 47px); }
  .subhero .lead { margin-top: 21px; font-size: 23px; }
  .subsection, .experience-section { padding-block: 43px; }
  .service-intro { font-size: 22px; }
  .project-grid { margin-top: 29px; }
  .project-item h3 { font-size: 25px; margin-bottom: 13px; }
  .project-item p { font-size: 19px; }
  .project-item { padding-top: 20px; }
  .output-grid .intro { margin-top: 18px; font-size: 20px; }
  .output-list li { font-size: 20px; padding-block: 17px; }
  .scope-note { font-size: 16px; margin-top: 20px; }
  .scope-questions { padding-left: 22px; }
  .scope-questions li { font-size: 20px; margin-bottom: 17px; padding-left: 5px; }
  .specialism p, .specialism li { font-size: 20px; }
  .related-services { flex-direction: column; align-items: start; gap: 6px; margin-top: 21px; }
  .about-page-text p { font-size: 21px; }
  .contact-page { padding-block: 24px 48px; }
  .contact-page h1 { font-size: clamp(39px, 9.4vw, 49px); }
  .contact-page .lead { font-size: 23px; margin-top: 22px; }
  .contact-email-block { margin-block: 31px; padding-block: 25px; gap: 22px; }
  .contact-email-block .email-link { font-size: clamp(21px, 5.4vw, 27px); }
  .contact-checklist { grid-template-columns: minmax(0, 1fr); gap: 26px; margin-top: 27px; }
  .contact-checklist h3 { font-size: 25px; margin-bottom: 12px; }
  .contact-checklist p { font-size: 20px; }
  .experience-intro { font-size: 22px; }
  .experience-list { margin-top: 28px; }
  .experience-row { grid-template-columns: minmax(0, 1fr); gap: 17px; padding-block: 24px 27px; }
  .experience-meta { font-size: 17px; }
  .experience-meta p, .experience-meta strong { margin-top: 6px; }
  .experience-detail h3 { margin-bottom: 15px; font-size: 27px; }
  .experience-detail p { font-size: 20px; }
  .publication-note { font-size: 18px; padding-top: 21px; }
  .histology-placeholder { margin-top: 24px; }
  .placeholder-art { min-height: 160px; max-height: 255px; }
  .histology-placeholder figcaption { padding: 13px 15px; }
  .work-item .placeholder-art { max-height: 245px; }
  .histomorph-intro { font-size: 22px; }
  .histomorph-feature { grid-template-columns: minmax(0, 1fr); gap: 28px; padding: 25px 23px; margin-block: 39px 49px; }
  .histomorph-feature p { font-size: 20px; }
  .histomorph-feature .placeholder-art { max-height: 230px; }
  .histomorph-grid { grid-template-columns: minmax(0, 1fr); gap: 32px; margin-top: 31px; }
  .histomorph-item { padding-top: 23px; }
  .histomorph-item h3 { font-size: 27px; }
  .histomorph-item p { font-size: 20px; }
  .histomorph-item .placeholder-art { max-height: 255px; }
  .measurement-list dd { font-size: 18px; }
  .sampling-note { margin-block: 39px; padding: 26px 23px; }
  .sampling-note p, .sampling-note li { font-size: 20px; }
  [id] { scroll-margin-top: 8px; }
  .tox-overview { font-size: 22px; }
  .toc-links { gap: 13px 25px; padding-block: 19px; }
  .toc-links a { font-size: 12px; }
  .tox-role-grid { grid-template-columns: minmax(0, 1fr); gap: 36px; padding-block: 42px; }
  .tox-role { padding-top: 23px; }
  .tox-role h2 { font-size: 34px; margin-bottom: 20px; }
  .tox-role p, .tox-role li { font-size: 20px; }
  .tox-role .histology-placeholder { margin-top: 24px; }
  .study-grid, .planning-grid, .quality-grid, .delivery-grid, .image-grid { grid-template-columns: minmax(0, 1fr); gap: 29px; }
  .study-grid { margin-top: 29px; }
  .study-grid h3 { font-size: 26px; }
  .planning-grid h2, .quality-grid h2, .delivery-grid h2 { margin-bottom: 20px; }
  .image-grid { margin-top: 29px; }
  .image-grid .placeholder-art { min-height: 170px; max-height: 255px; }
  .quality-note { font-size: 17px; margin-top: 22px; }
  .portrait-placeholder { max-width: 240px; }
  html:not(.js) .menu-toggle { display: none; }
  html:not(.js) .header-inner { flex-wrap: wrap; gap: 13px; padding-block: 14px; }
  html:not(.js) .nav { position: static; display: flex; flex-direction: row; flex-wrap: wrap; width: 100%; gap: 7px 20px; padding: 0; border: 0; }
  html:not(.js) .nav > a:not(.button) { padding-block: 8px; }
  html:not(.js) .nav .button { margin: 0; }
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { animation: none !important; transition: none !important; }
}

/* More space for images, with practical detail available on request. */
.hero-illustrated {
  display: grid;
  grid-template-columns: minmax(0, 1.12fr) minmax(0, .88fr);
  align-items: center;
  gap: 70px;
  padding-block: 62px 72px;
}
.hero-illustrated > * { min-width: 0; }
.hero-illustrated .hero-text > .eyebrow { margin-bottom: 24px; }
.hero-illustrated h1 { max-width: 660px; font-size: clamp(48px, 4.8vw, 64px); }
.hero-illustrated .hero-copy { margin-top: 27px; font-size: 22px; max-width: 560px; }
.hero-background { margin-top: 25px; color: var(--muted); font-size: 15px; line-height: 1.5; }
.hero-background a { color: var(--forest); text-decoration: underline; text-underline-offset: 4px; }
.hero-flexibility { margin-top: 17px; max-width: 560px; color: var(--forest); font-size: 17px; line-height: 1.55; }
.hero-visual { min-width: 0; padding: 24px; background: var(--lagoon-wash); border-top: 3px solid var(--lagoon); }
.hero-visual .histology-placeholder { max-width: none; width: 100%; margin: 0; background: var(--paper); }
.hero-visual .placeholder-art { min-height: 310px; max-height: 460px; aspect-ratio: 1 / 1; background: #d5e9e3; }
.work-note > .histology-placeholder { width: 100%; max-width: none; margin: 0 0 24px; }
.work-note .placeholder-art { min-height: 190px; max-height: 270px; aspect-ratio: 16 / 9; }
.work-note:nth-child(2) .placeholder-art { background: var(--coral-wash); color: #805447; }
.personal-meta .portrait-placeholder { max-width: 280px; margin-top: 25px; }
.personal-meta .portrait-placeholder .placeholder-art { min-height: 220px; max-height: 320px; }
.model-showcase { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 30px; }
.model-example { min-width: 0; }
.model-example .histology-placeholder { max-width: none; margin: 0 0 22px; }
.model-example .placeholder-art { min-height: 190px; max-height: 250px; }
.model-example:nth-child(2) .placeholder-art { background: #e1e9dd; }
.model-example:nth-child(3) .placeholder-art { background: var(--coral-wash); color: #805447; }
.model-example h3 { font-size: 25px; margin-bottom: 13px; }
.model-example > p { color: var(--muted); font-size: 18px; }
.histomorph-item .histology-placeholder { margin: 0 0 22px; width: 100%; max-width: none; }
.histomorph-item .placeholder-art { min-height: 230px; max-height: 310px; aspect-ratio: 16 / 10; }
.histomorph-item .measurement-summary { font-size: 19px; line-height: 1.55; }
.method-detail, .assignment-detail { margin-top: 23px; border-block: 1px solid var(--line); }
.method-detail > summary, .assignment-detail > summary {
  display: flex;
  justify-content: space-between;
  align-items: baseline;
  gap: 20px;
  padding-block: 16px;
  color: var(--forest);
  cursor: pointer;
  list-style: none;
  font-size: 17px;
  line-height: 1.45;
}
.method-detail > summary::-webkit-details-marker, .assignment-detail > summary::-webkit-details-marker { display: none; }
.method-detail > summary::after, .assignment-detail > summary::after { content: '+'; flex: 0 0 auto; font-size: 21px; line-height: 1; color: var(--lagoon-text); }
.method-detail[open] > summary::after, .assignment-detail[open] > summary::after { content: '−'; }
.method-detail > summary:hover, .assignment-detail > summary:hover { color: var(--lagoon-text); }
.method-detail > p { padding-bottom: 20px; color: var(--muted); font-size: 17px; line-height: 1.6; }
.assignment-detail .measurement-list { margin: 0 0 6px; }
.assignment-detail .measurement-list > div:first-child { border-top: 0; padding-top: 0; }
.assignment-detail .measurement-list dd { font-size: 17px; }
.engagement-deliverable { padding-top: 4px; color: var(--muted); font-size: 19px; line-height: 1.55; }
.engagement-deliverable .eyebrow { display: block; margin-bottom: 12px; }
.engagement-details > .button { margin-top: 24px; }
.oncology-visuals { align-items: center; }
.oncology-visuals > .histology-placeholder { width: 100%; max-width: 430px; }
.oncology-visuals .placeholder-art { min-height: 280px; max-height: 430px; aspect-ratio: 1 / 1; }
.about-page-text .career-list { margin-top: 28px; }

@media (max-width: 1050px) {
  .hero-illustrated { gap: 40px; }
  .hero-visual { padding: 18px; }
  .hero-visual .placeholder-art { min-height: 280px; }
}
@media (max-width: 900px) {
  .hero-illustrated { grid-template-columns: minmax(0, 1fr); gap: 38px; }
  .hero-illustrated h1 { max-width: 720px; font-size: 55px; }
  .hero-illustrated .hero-copy { max-width: 650px; }
  .hero-visual { width: 100%; padding: 24px; }
  .hero-visual .placeholder-art { min-height: 250px; max-height: 340px; aspect-ratio: 16 / 9; }
  .model-showcase { grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 30px; }
}
@media (max-width: 680px) {
  .hero-illustrated { padding-block: 39px 44px; gap: 31px; }
  .hero-illustrated .hero-text > .eyebrow { margin-bottom: 21px; }
  .hero-illustrated h1 { font-size: clamp(38px, 9vw, 45px); line-height: 1.1; }
  .hero-illustrated .hero-copy { margin-top: 23px; font-size: 21px; }
  .hero-background { margin-top: 22px; font-size: 14px; }
  .hero-visual { padding: 16px; }
  .hero-visual .placeholder-art { min-height: 220px; max-height: 300px; aspect-ratio: 4 / 3; }
  .work-note > .histology-placeholder { margin-bottom: 21px; }
  .work-note .placeholder-art { min-height: 190px; max-height: 250px; }
  .personal-meta .personal-portrait { margin-top: 23px; }
  .model-showcase { grid-template-columns: minmax(0, 1fr); gap: 33px; }
  .model-example .placeholder-art { min-height: 200px; max-height: 270px; }
  .model-example > p { font-size: 19px; }
  .histomorph-item .placeholder-art { min-height: 210px; max-height: 280px; }
  .histomorph-item .measurement-summary { font-size: 19px; }
  .method-detail > p, .assignment-detail .measurement-list dd { font-size: 17px; }
  .oncology-visuals .placeholder-art { min-height: 240px; max-height: 330px; }
  .about-page-grid .personal-portrait { max-width: 280px; }
}

/* Original concept illustrations retain their complete landscape compositions. */
.editorial-illustration {
  width: 100%;
  max-width: 100%;
  min-width: 0;
  margin: 0;
  border: 0;
  background: var(--paper);
}
.editorial-illustration img {
  display: block;
  width: 100%;
  height: auto;
  aspect-ratio: 3 / 2;
  object-fit: contain;
}
.editorial-illustration figcaption {
  padding: 12px 0 0;
  border-top: 1px solid var(--line);
  color: var(--muted);
  font-size: 13px;
  line-height: 1.5;
  overflow-wrap: anywhere;
}
.illustration-label {
  display: block;
  margin-bottom: 3px;
  color: var(--lagoon-text);
  font: 600 11px/1.45 var(--nav);
}
.hero-visual > .editorial-illustration,
.featured-work > .editorial-illustration,
.planning-grid > .editorial-illustration { max-width: none; }
.tox-role > .editorial-illustration { margin-top: auto; }
.histomorph-item > .editorial-illustration { margin: 0 0 22px; }
.illustrated-intro { display: grid; grid-template-columns: minmax(0, 1.1fr) minmax(0, .9fr); gap: 58px; align-items: center; }
.illustrated-intro > * { min-width: 0; }
.illustrated-intro + .toc-links { margin-top: 35px; }
.application-range h3 { margin-top: 20px; font-size: 20px; }
.application-range p { max-width: 940px; }
.name-story-reflection { margin-top: 17px; }
@media (max-width: 820px) {
  .illustrated-intro { grid-template-columns: minmax(0, 1fr); gap: 27px; }
  .illustrated-intro > .editorial-illustration { max-width: 540px; margin-inline: auto; }
  .tox-role > .editorial-illustration { margin-top: 12px; }
}
.work-note > .editorial-illustration { margin-bottom: 24px; }
.model-example > .editorial-illustration { margin-bottom: 22px; }

/* Model applications: an illustrated lead, then a readable two-column set. */
.model-page-intro { padding-bottom: 38px; }
.model-page-note { max-width: 770px; margin-top: 24px; color: var(--muted); font-size: 17px; }
.model-page-note a { color: var(--lagoon-text); text-decoration: underline; text-underline-offset: 4px; }
.model-area-list { max-width: 1080px; margin-top: 25px; color: var(--muted); font-size: 18px; line-height: 1.75; }
.model-overview-intro { max-width: 850px; margin-bottom: 30px; color: var(--muted); font-size: 18px; }
.model-overview-grid { display: grid; grid-template-columns: minmax(0, .9fr) minmax(0, 1.1fr); gap: 56px; align-items: start; }
.model-index { min-width: 0; }
.model-index-link { display: grid; grid-template-columns: minmax(0, 1fr) 20px; gap: 14px; padding: 16px 0; border-top: 1px solid var(--line); }
.model-index-link strong { display: block; font-size: 20px; font-weight: 400; color: var(--forest); }
.model-index-link > span > span { display: block; margin-top: 3px; font-size: 15px; color: var(--muted); }
.model-index-link svg { width: 20px; height: 20px; margin-top: 4px; color: var(--lagoon-text); }
.model-index-link:hover strong { color: var(--lagoon-text); }
.model-focus-grid { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 50px 56px; padding-bottom: 60px; }
.model-focus-panel { min-width: 0; padding-top: 24px; border-top: 2px solid var(--lagoon); }
.model-focus-panel:nth-child(2n) { border-top-color: var(--forest); }
.model-focus-panel:nth-child(4) { border-top-color: var(--coral); }
.model-focus-copy { min-width: 0; padding-top: 24px; }
.model-focus-copy h2 { margin-top: 10px; font-size: 31px; line-height: 1.15; }
.model-examples { margin-top: 15px; color: var(--lagoon-text); font-size: 15px; line-height: 1.5; }
.model-summary { margin-top: 18px; color: var(--muted); font-size: 19px; }
.model-focus-lead { grid-column: 1 / -1; display: grid; grid-template-columns: minmax(0, 1fr) minmax(0, 1fr); gap: 56px; padding-bottom: 10px; }
.model-focus-lead .model-focus-copy { padding-top: 5px; }
.model-focus-lead h2 { font-size: 38px; }
.model-method { margin-top: 22px; }
.model-method > summary { font-size: 16px; }
.model-method dl > div { padding-bottom: 18px; }
.model-method dt { margin-bottom: 5px; color: var(--forest); font-size: 15px; font-weight: 600; }
.model-method dd { color: var(--muted); font-size: 17px; line-height: 1.6; }
.model-references { padding-bottom: 20px; font-size: 13px; }
.model-references p { color: var(--forest); margin-bottom: 5px; }
.model-references a { display: block; padding-block: 4px; color: var(--lagoon-text); text-decoration: underline; text-underline-offset: 3px; }
.model-planning { display: grid; grid-template-columns: minmax(0, .7fr) minmax(0, 1.3fr); gap: 56px; }
.model-planning h2 { margin-top: 12px; font-size: 32px; }

/* Image analysis keeps the application range open and method detail optional. */
.analysis-applications { padding-top: 40px; }
.analysis-applications .section-heading { max-width: 800px; }
.analysis-breadth-grid { display: grid; grid-template-columns: minmax(0, 1.3fr) minmax(0, 1fr); gap: 56px; }
.analysis-breadth-grid h3 { font-size: 23px; margin-bottom: 15px; }
.analysis-breadth-grid p { color: var(--muted); font-size: 18px; line-height: 1.65; }
.analysis-methods-intro { margin-top: 28px; max-width: 1000px; font-size: 19px; color: var(--muted); }
.analysis-examples { padding-top: 15px; }
.analysis-focus-panel h3 { margin-top: 10px; font-size: 31px; line-height: 1.15; }
.analysis-focus-panel.model-focus-lead h3 { font-size: 38px; }
.analysis-context { margin-top: 17px; color: var(--muted); font-size: 17px; line-height: 1.6; }
.analysis-workflow-grid { display: grid; grid-template-columns: minmax(0, 1fr) minmax(0, 1fr); gap: 56px; }
.analysis-workflow-grid h2 { margin-block: 15px 24px; }
.analysis-workflow-grid p { color: var(--muted); font-size: 19px; }
.analysis-inputs { padding-left: 22px; margin: 0 0 20px; color: var(--muted); font-size: 17px; }
.analysis-inputs li + li { margin-top: 15px; }
.tox-scope-line { margin-top: 20px; color: var(--lagoon-text); font-size: 16px; }
.scientific-work-list { display: grid; grid-template-columns: repeat(2,minmax(0,1fr)); gap: 42px; margin-top: 30px; }
.scientific-work-list article { min-width: 0; padding-top: 21px; border-top: 2px solid var(--lagoon); }
.scientific-work-list h3 { margin-top: 12px; font-size: 24px; line-height: 1.3; }
.scientific-work-list .work-role { margin-top: 18px; color: var(--muted); font-size: 16px; }
.scientific-work-note { margin-top: 27px; color: var(--muted); font-size: 15px; }
.contact-tools { display: flex; align-items: center; flex-wrap: wrap; gap: 10px 24px; margin-top: 18px; }
.contact-tools .text-link { min-height: 44px; }
.copy-status { color: var(--muted); font-size: 15px; }
.enquiry-faq { margin-top: 45px; }
.enquiry-faq h2 { margin-bottom: 23px; }
.enquiry-faq details + details { border-top: 0; margin-top: 0; }
.enquiry-faq details { max-width: 900px; margin-top: 0; }
.toc-links a { display: inline-flex; align-items: center; min-height: 44px; padding-block: 7px; font-size: 14px; }

@media (max-width: 900px) {
  .analysis-breadth-grid, .analysis-workflow-grid { gap: 30px; }
  .analysis-focus-panel.model-focus-lead h3 { font-size: 32px; }
  .model-overview-grid { gap: 30px; }
  .model-focus-grid { gap: 40px 30px; }
  .model-focus-lead { gap: 30px; }
  .model-focus-lead h2 { font-size: 32px; }
  .model-planning { gap: 30px; }
}
@media (max-width: 680px) {
  .nav-services { width: 100%; font-size: 15px; }
  .nav.is-open { max-height: calc(100vh - 100px); max-height: calc(100dvh - 100px); overflow-y: auto; overscroll-behavior: contain; }
  .nav-services > summary { padding-block: 13px; }
  .service-menu { position: static; width: 100%; padding: 2px 0 10px 10px; border: 0; border-left: 2px solid var(--line); box-shadow: none; }
  .service-menu a { padding: 11px 12px; }
  .scientific-work-list { grid-template-columns: minmax(0,1fr); gap: 28px; }
  .scientific-work-list h3 { font-size: 24px; }
  .toc-links a { font-size: 13px; }
  .analysis-breadth-grid, .analysis-workflow-grid { grid-template-columns: minmax(0, 1fr); gap: 28px; }
  .analysis-applications { padding-top: 31px; }
  .analysis-focus-panel h3, .analysis-focus-panel.model-focus-lead h3 { font-size: 30px; }
  .model-page-intro { padding-bottom: 26px; }
  .model-overview-grid, .model-focus-grid, .model-focus-lead, .model-planning { grid-template-columns: minmax(0, 1fr); }
  .model-focus-grid { gap: 37px; padding-bottom: 38px; }
  .model-focus-lead { gap: 0; }
  .model-focus-lead .model-focus-copy { padding-top: 24px; }
  .model-focus-copy h2, .model-focus-lead h2 { font-size: 30px; }
  .model-overview-grid { gap: 25px; }
  .model-index-link { padding-block: 15px; }
  .model-planning { gap: 22px; }
}

@media (max-width: 680px) {
  .editorial-illustration figcaption { padding-top: 10px; }
  .work-note > .editorial-illustration { margin-bottom: 21px; }
}